1. INRA, UMR1202 Biodiversity, Genes and Communities, 69 Route d'Arcachon, F-33612 Cestas, France.
2. University of Bordeaux I, UMR1202 Biodiversity, Genes and Communities, 69 Route d'Arcachon, F-33612 Cestas, France.
3. Departments of Plant Biology and Geology, and Program in Ecology, Evolution, and Conservation Biology, University of Illinois, Urbana, IL 61801, USA.
4. Department of Ecology and Evolutionary Biology and Herbarium, University of Michigan, 830 North University Avenue, Ann Arbor, MI 48103, USA.
5. Smithsonian Tropical Research Institute, Post Office Box 0843-03092, Balboa, Ancón, Republic of Panama.