1. Biology Department, Unit of Ecology and Systematics, Vrije Universiteit Brussel, Pleinlaan 2, 1050 Brussels, Belgium.
2. Laboratory of Evolutionary Genetics, Université Libre de Bruxelles, Code Postal, 300, Institute for Molecular Biology and Medicine, Rue Jeener and Brachet 12, B-6041 Gosselies, Belgium.
3. Department of Biology, Boston University, 5 Cummington Street, Boston, MA 02215, USA.
4. Wildlife Heritage Trust, 95 Cotta Road, Colombo 8, Sri Lanka.
5. Department of Zoology, The Natural History Museum, London SW7 5BD, UK.