1. Department of Natural History Sciences, Faculty of Science, Hokkaido University, Sapporo 060-0810, Japan
2. Department of Botany, 3190 Maile Way, University of Hawaii at Manoa, Honolulu, Hawaii 96822, USA
3. Non-Vascular Plants Research Division, National Institute of Biological Resources, Environmental Research Complex, Gyeongseo-dong, Seo-gu, Incheon 404-708, Korea
4. Department of Biology, Chungnam National University, Daejon 305-764, Korea