1. Shimoda Marine Research Centre, University of Tsukuba, 5-10-1 Shimoda, Shizuoka 415-0025, Japan, and School of Life & Environmental Sciences, Deakin University, PO Box 423, Warrnambool, Victoria 3280, Australia
2. Shimoda Marine Research Centre, University of Tsukuba, 5-10-1 Shimoda, Shizuoka 415-0025, Japan