1. Graduate School of Kuroshio Science, Kochi University, 200 Otsu, Monobe, Nankoku, Kochi 783-8502, Japan
2. Faculty of Core Research, Natural Science Division, Ochanomizu University, 2-1-1 Otsuka, Bunkyo, Tokyo 112-8610, Japan
3. Muroran Marine Station, Field Science Center for Northern Biosphere, Hokkaido University, 1-133-31, Funami-cho, Muroran, Hokkaido 051-0013, Japan
4. Research and Education Faculty, Natural Sciences Cluster, Sciences Unit, Kochi University, 2-5-1 Akebono, Kochi 780-8520, Japan
5. College of Marine Ecology and Environment, Shanghai Ocean University, Shanghai 201306, China
6. Usa Marine Biological Institute, Kochi University, Inoshiri 194, Usa, Tosa, Kochi 781-1164, Japan