1. Division of Environmental Biotechnology, Graduate School of Global Environmental Studies, Kyoto University, Yoshida-Honmachi, Sakyo-ku, Kyoto 606-8501, Japan
2. Division of Marine Bioresource and Environmental Science, Graduate School of Fisheries Sciences, Hokkaido University, Minato-cho 3-3-1, Hakodate, Hokkaido 041-8611, Japan
3. Division of Applied Biosciences, Graduate School of Agriculture, Kyoto University, Kitashirakawa-Oiwake-cho, Sakyo-ku, Kyoto 606-8502, Japan
4. Division of Forest and Biomaterials Science, Graduate School of Agriculture, Kyoto University, Kitashirakawa-Oiwake-cho, Kyoto 606-8502, Japan
5. Faculty of Bioresources, Mie University, 1577 Kurima-machiya-cho, Tsu, Mie 514-8507, Japan