1. Faculty of Biotechnology, Fukui Prefectural University, 1-1 Gakuencho, Obama, Fukui, 917-0003, Japan
2. School of Biosciences 2, University of Melbourne, Parkville Victoria 3010, Australia
3. Institute of Biological Sciences, Applied Ecology & Phycology, University of Rostock, D-18059 Rostock, Germany
4. Instituto Oceanográfico, Universidad de Oriente, Cumaná 6101, Venezuela