1. Guangdong Provincial Key Laboratory of Healthy and Safe Aquaculture, College of Life Science, South China Normal University, West 55 of Zhongshan Road, Guangzhou 510631, P.R. China
2. Natural History Museum of Denmark, University of Copenhagen, Sølvgade 83S, 1307 Copenhagen K, Denmark
3. Section of Marine Biology, Institute of Biology, University of Copenhagen, Universitetsparken 4, DK-2100 Copenhagen Ø, Denmark