1. Third Institute of Oceanography, State Oceanic Administration, Xiamen 361005, China
2. Ifremer, LER BO, Station de Biologie Marine, Place de la Croix, BP40537, F-29185 Concarneau Cedex, France
3. Stazione Zoologica Anton Dohrn, Department of Integrative Marine Ecology, Villa Comunale, 80121 Naples, Italy
4. CNRS, UMR7144, Station Biologique de Roscoff, 29680, Roscoff, France
5. Ifremer – Centre de Brest, DYNECO PELAGOS, F-29280 Plouzané, France