1. College of Marine Ecology and Environment, Shanghai Ocean University, Shanghai 201306, China
2. Marine Resources Development Institute of Jiangsu, Huaihai Institute of Technology, Lianyungang 222005, China
3. Department of Marine Science, College of Natural Sciences, Incheon National University, Incheon 22012, Korea
4. Department of Ecology and Evolutionary Biology, University of Connecticut, 1 University Place, Stamford, CT 06901, USA
5. Research Institute of Basic Sciences, Incheon National University, Incheon 22012, Korea