1. Laboratory ‘Terre Océan’, University of French Polynesia, B. P. 6570 Faa'a 98702, Tahiti, French Polynesia
2. UMR 7138 ‘Systematics, Adaptation, Evolution’ IRD-Noumea - BPA5, 98848 Nouméa cedex, New Caledonia
3. Herbarium, University of North Carolina, Chapel Hill, NC 27599-3280, USA
4. Department of Biology, University of Louisiana at Lafayette, Lafayette, LA 70504-2451, USA