1. Graduate School of Natural Sciences, Kobe University, Rokkodai, Kobe 657-8501, Japan
2. Department of Plant Biology, Faculty of Science, University of Malaga, Campus de Teatinos s/n, E-29071 Malaga, Spain
3. Department of Botany and Plant Pathology, Oregon State University, Cordley Hall 2082, Corvallis, OR 97331-2902
4. Faculty of Biology, University of Konstanz, D-78434 Konstanz, Germany
5. Kobe University Research Center for Inland Seas, Rokkodai, Kobe 657-8501, Japan