1. Graduate School of Agricultural and Life Sciences, University of Tokyo, 1-1-1 Yayoi, Bunkyo-ku, Tokyo 113-8657, Japan
2. Marine Biological Research Institute, Zoology Department, University of Cape Town, Rondebosch, Cape Town 7701, South Africa
3. Japan NUS Co. Ltd, 1-1 Fukuura, Kanazawa-ku, Yokohama 236-0004, Japan
4. Iwate Fisheries Technology Center, 3-75-3 Heita, Kamaishi, Iwate 026-0001, Japan
5. Asian Natural Environmental Science Center, University of Tokyo, 1-1-1 Yayoi, Bunkyo-ku, Tokyo 113-8657, Japan