1. Graduate School of Natural Science, Konan University, Okamoto, Higashinada, Kobe, Hyogo 658-8501, Japan
2. Department of Biology, Kongju National University, Kongju 314-701, Korea
3. School of Botany, University of Melbourne, Parkville, Victoria 3010, Australia
4. Division of Biology, Newcastle University, Newcastle upon Tyne NE1 7RU, UK
5. Department of Biology, Faculty of Science and Engineering, Konan University, Okamoto, Higashinada, Kobe, Hyogo 658-8501, Japan