1. United Graduate School of Agricultural Sciences, Kagoshima University, Korimoto 1-21-24, Kagoshima 890-0065, Japan
2. Faculty of Fisheries, Kagoshima University, Shimoarata 4-50-20, Kagoshima 890-0056, Japan
3. Institute for East China Sea Research, Organisation for Marine Science and Technology, Nagasaki University, Taira-machi 1557-7, Nagasaki 851-2213, Japan
4. Faculty of Core Research, Natural Science Division, Ochanomizu University, 2-1-1 Otsuka, Bunkyo, Tokyo 112-8610, Japan