1. Muroran Marine Station, Field Science Center for Northern Biosphere, Hokkaido University, Muroran 051-0013, Japan
2. Faculty of Marine Bioscience, Fukui Prefectural University, Obama 917-0003, Japan
3. Faculty of Environmental & Symbiotic Sciences, Prefectural University of Kumamoto, Kumamoto 862-8502, Japan
4. Tohoku National Fisheries Research Institute, Fisheries Research Agency, Shiogama 985-0001, Japan