1. Hebei Key Laboratory of Ocean Dynamics, Resources and Environments, Hebei Normal University of Science and Technology, Qinhuangdao, 066004, China
2. College of Marine Resources and Environment, Hebei Normal University of Science and Technology, Qinhuangdao, 066004, China
3. Marine Living Resources and Environment Key Laboratory of Hebei Province, Qinhuangdao, 066004, China
4. Guangxi Key Laboratory of Beibu Gulf Marine Biodiversity Conservation, College of Marine Sciences, Beibu Gulf University, Qinzhou, 535011, China