1. South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences/Key Lab. of South China Sea Fishery Resources Exploitation and Utilization, Ministry of Agriculture and Rural Affairs, Guangzhou, 510300, P.R. China; Tropical Fishery Research and Development Center, South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Sanya 572018, P.R. China
2. South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences/Key Lab. of South China Sea Fishery Resources Exploitation and Utilization, Ministry of Agriculture and Rural Affairs, Guangzhou, 510300, P.R. China
3. South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences/Key Lab. of South China Sea Fishery Resources Exploitation and Utilization, Ministry of Agriculture and Rural Affairs, Guangzhou, 510300, P.R. China;