1. College of Fisheries and Life Sciences, Shanghai Ocean University, Shanghai 201306, , P.R. China
2. Zhuhai modern agricultural development center, 519070, Zhuhai, Guangdong Province, P.R. China
3. Key Laboratory of South China Sea Fishery Resources Exploitation and Utilization, Ministry of Agriculture and Rural Affairs; South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, 510300, Guangzhou, Guangdong Province, P.R. China