1. Marine Science Research Institute of Shandong Province (National Oceanographic Center), Qingdao Shandong 266104, China
2. Shandong Key Laboratory of Disease Control in Mariculture, Qingdao Shandong 266104, China
3. National Pathogen Collection Center for Aquatic Animals, Shanghai Ocean University, Shanghai 201306, China
4. Key Laboratory of Freshwater Fishery Germplasm Resources, Ministry of Agriculture and Rural Affairs of China, Shanghai 201306, China