1. School of Biological Sciences, Royal Holloway, University of London, Egham, TW20 0EX, UK
2. Department of Biology and Ecology of Fishes, Leibniz-Institute of Freshwater Ecology and Inland Fisheries, Müggelseedamm 310, 12587 Berlin, Germany
3. Department Physiological Chemistry, Biocenter, University of Würzburg, and Comprehensive Cancer Center Mainfranken, University Clinic Würzburg, 97078 Würzburg, Germany, and Hagler Institute for Advanced Studies and Department of Biology, Texas A&M Un