1. Echigo-Matsunoyama Museum of Natural Sicences, ‘Kyororo’, 1712-2 Matsunoyama, Tôkamachi, 942-1411, Japan
2. Biosystematics Laboratory, Faculty of Social and Cultural Studies, Kyushu University, 744 Motooka, Fukuoka 819-0395, Japan
3. The Shirakami Institute for Environmental Sciences, Hirosaki University, Bunkyo-cho 3, Hirosaki City, Aomori 036-8561, Japan