1. National Agricultural Research Center, NARO, Tsukuba, Ibaraki 305-8666, Japan
2. National Agricultural Research Center for Tohoku Region, NARO, Fukushima, Fukushima 960-2156, Japan
3. National Institute of Agrobiological Sciences, Tsukuba, Ibaraki 305-8602, Japan
4. Faculty of Agriculture, Tohoku University, Aoba-ku, Sendai, Miyagi 981-8555, Japan