1. The United Graduate School of Agricultural Science, Ehime University, Ko 498, Hattanji, Matsuyama, Ehime 799-2424, Japan
2. National Agricultural Research Center for Tohoku region, Yotsuya, Daisen, Akita 014-0102, Japan
3. National Agricultural Research Center, Kannondai, Tsukuba, Ibaraki 305-8666, Japan