1. Université Lyon, Université Claude Bernard Lyon 1, ENS Lyon, CNRS, UMR 5276 Laboratoire de Géologie de Lyon: Terre, Planètes, Environnement, F-69622 Villeurbanne, France
2. Department of Natural Sciences, National Museums Scotland, Edinburgh EH1 1JF, UK
3. Palaeoecosystems Group, Department of Earth Sciences, Durham University, Durham DH1 3LE, UK
4. Czech Geological Survey; Klárov 3, Prague 1, 118 21, Czech Republic