1. Deparment of Botany, University of Wyoming, 1000 E. University Ave., Laramie, Wyoming, 82071, USA
2. Department of Geology and Geophysics, University of Wyoming, 1000 E. University Ave, Laramie, Wyoming 82071, USA
3. Department of Geosciences, University of Arizona, 1040 E. 4th St., Tucson, Arizona 85721, USA