1. State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Beijing 100083, China
2. School of the Earth Sciences and Resources, China University of Geosciences, Beijing 100083, China
3. Dinosaur Trackers Research Group, University of Colorado, Denver, Colorado 80217, USA
4. Mace Brown Museum of Natural History, Department of Geology and Environmental Geosciences, College of Charleston, Charleston, South Carolina 29401, USA
5. Saurierwelt Paläontologisches Museum, Neumarkt D-92318, Germany
6. School of Biological Sciences, the University of Queensland, Queensland 4072, Australia