1. Centre for Bioethics and Risk Assessment, Division of Laboratory Animal Science and Welfare, Department of Veterinary Pathobiology, Faculty of Life Sciences, University of Copenhagen, Groennegaardsvej 15, DK-1870 Frederiksberg C, Denmark
2. Division of Veterinary Epidemiology, Department of Large Animal Sciences, Faculty of Life Sciences, University of Copenhagen, Groennegaardsvej 8, DK-1870 Frederiksberg C, Denmark
3. Institute of Oral Medicine, Dental School, University of Copenhagen, Nørre Allé 20, DK-2200 Copenhagen N, Denmark
4. Immunology Research Centre, St Vincents Hospital, 41 Victoria Parade, Fitzroy, Victoria 3065, Australia