1. Instituto Gulbenkian de Ciência 1 , Oeiras 2780-156 , Portugal
2. Åbo Akademi University 2 Faculty of Science and Engineering, Cell Biology , , Turku 20500 , Finland
3. Turku Bioscience Centre 3 , , Turku 20520 , Finland
4. University of Turku and Åbo Akademi University 3 , , Turku 20520 , Finland
5. University of Turku and Åbo Akademi University 4 Turku Bioimaging , , Turku 20520 , Finland
6. Åbo Akademi University 5 InFLAMES Research Flagship Center , , Turku 20100 , Finland
7. University College London 6 UCL Laboratory for Molecular Cell Biology , , London WC1E 6BT , UK