1. Applied Vision Research Centre, The Henry Wellcome Laboratories for Vision Sciences, Department of Optometry and Visual Science, City University,Northampton Square, London EC1V 0HB, UK
2. School of Biological Sciences, University of Plymouth, Drake Circus,Plymouth PL4 8AA, UK
3. Anatomisches Institut, Universität Tübingen,Österbergstrasse 3, 72074 Tübingen, Germany