1. Department of Organismal Biology and Anatomy, University of Chicago, Chicago, IL 60637, USA
2. Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada T6G 2E9
3. Department of Dentistry, University of Alberta, Edmonton, Alberta, Canada T6G 2R3
4. Department of Stem Cell Biology and Regenerative Medicine, W.M. Keck School of Medicine, University of Southern California, Los Angeles, CA 90033, USA
5. Developmental Biology Unit, European Molecular Biology Laboratory, Heidelberg, 69117, Germany
6. Department of Medical Genetics, University of Alberta, Edmonton, Alberta, Canada T6G 2R7