1. Stowers Institute for Medical Research, Kansas City, MO 64110, USA
2. Departments of Orofacial Sciences and Pediatrics, Program in Craniofacial and Mesenchymal Biology, University of California at San Francisco, San Francisco, CA 94143-0442, USA
3. Department of Anatomy and Cell Biology, University of Kansas Medical Center, Kansas City, KS 66160, USA