1. Norwegian University of Science and Technology 1 Department of Biology , , Trondheim NO-7491 , Norway
2. Research and Consultancy van der Kooij 2 Nature Education , , Rudsteinveien 67, Slattum NO-1480 , Norway
3. Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences 3 , Box 5003, Ås NO-1433 , Norway
4. School of Science, Technology and Engineering, University of the Sunshine Coast 4 , Maroochydore DC, QLD 4558 , Australia