1. University of Cincinnati 1 Department of Biological Sciences , , Cincinnati, OH 45221 , USA
2. College of Environmental Science and Forestry, State University of New York 2 Department of Environmental Biology , , Syracuse, NY 13210 , USA
3. Emory University 3 Department of Biology , , Atlanta, GA 30322 , USA
4. University of Kansas 4 Department of Ecology and Evolutionary Biology , , Lawrence, KS 66045 , USA