1. Institut Curie, PSL Research University, CNRS UMR3348, F-91405 Orsay, France
2. Université Paris Sud, Université Paris-Saclay, CNRS UMR3348, F-91405 Orsay, France
3. current address: Institut Pasteur, Laboratory of Genetics and Physiology of Hearing, INSERM UMRS1120, F-75724 Paris, France
4. CERTO Centre d'Etudes et de Recherches Thérapeutiques en Ophtalmologie, Université Paris Sud, Université Paris-Saclay, CNRS UMR9197, F-91405 Orsay, France
5. current address: McGill University, Department of Biology, Montréal, Québec H3A 1B1, Canada
6. current address: Institut d'Investigacions Biomèdiques August Pi i Sunyer (IDIBAPS), Hospital Clínic de Barcelona, E-08036 Barcelona, Spain
7. current address: Harvard Medical School, Children's Hospital Boston, Boston MA 02115, USA
8. Institut Curie, PSL Research University, Department of Genetics, F-75005, Paris, France
9. Université Paris Descartes, Sorbonne Paris Cité, F-75005, Paris, France