1. Cambridge University Department of Physiology Development and Neuroscience, Downing St, Cambridge, CB2 3EG, UK
2. Program in Cell Biology, The Hospital for Sick Children, Peter Gilgan Centre for Research and Learning, 686 Bay Street, Room 15.9716, Toronto, ON M5G 0A4, Canada
3. Department of Molecular Genetics, University of Toronto, Room 4396, Medical Sciences Building, 1 King's College Circle, Toronto, ON M5S 1A8, Canada