1. George Mason University 1 Department of Biology , , Fairfax, VA 22030 , USA
2. Davidson College 2 Department of Biology , , Davidson, NC 28035 , USA
3. School of Biological Sciences, Illinois State University 3 Behavior, Ecology, Evolution, and Conservation Section , , Normal, IL 61790 , USA
4. School of Integrative Biology, University of Illinois Urbana-Champaign 4 Department of Evolution, Ecology, and Behavior , , Urbana, IL 61801 , USA