1. University of Alberta 1 Department of Cell Biology, Faculty of Medicine and Dentistry , , Edmonton, Alberta , Canada
2. Universidad de la República, 11400 2 Sección Biología Celular, Facultad de Ciencias , , Uruguay
3. Institut Pasteur Montevideo 3 , Uruguay
4. University of Alberta 4 Division of Anatomy, Department of Surgery, Faculty of Medicine and Dentistry , , Edmonton, Alberta T6G 2H7 , Canada
5. University of Alberta 5 Department of Medical Genetics, Faculty of Medicine and Dentistry , , Edmonton, Alberta , Canada
6. Women and Children's Health Research Institute, University of Alberta 6 , Edmonton, Alberta , Canada