1. The Ohio State University Medical Center 1 Department of Biological Chemistry and Pharmacology , , Columbus, OH 43210 , USA
2. The Ohio State University 2 Department of Molecular Genetics , , Columbus, OH 43210 , USA
3. Center for RNA Molecular Biology 3 , Department of Biochemistry, School of Medicine , , Cleveland, OH 44106 USA
4. Case Western Reserve University 3 , Department of Biochemistry, School of Medicine , , Cleveland, OH 44106 USA
5. University of Wisconsin-Madison 4 Department of Cell and Regenerative Biology , , Madison, WI 53705 , USA
6. The Ohio State University 5 Department of Evolution, Ecology, and Organismal Biology , , Columbus, OH 43210 , USA
7. Department of Cell Biology, Duke Regeneration Center, Duke University School of Medicine 6 , Durham, NC 27710 , USA
8. The Ohio State University Medical Center 7 Department of Cell Biology and Physiology , , Columbus, OH 43210 , USA