1. Scripps Institution of Oceanography, University of California San Diego, Center for Marine Biotechnology and Biomedicine, 8655 Kennel Way, La Jolla, CA 92037, USA
2. Current address: National Marine Mammal Foundation, 2240 Shelter Island Drive, Suite 200, San Diego, California 92106, USA
3. Atmosphere and Ocean Research Institute, The University of Tokyo, 5-1-5 Kashiwanoha, Kashiwa, Chiba 277-8564, Japan