1. School of Immunology and Microbial Sciences, King's College London 1 Department of Infectious Diseases , , SE1 9RT London , UK
2. Wellcome-Wolfson Institute for Experimental Medicine, School of Medicine, Dentistry and Biomedical Sciences, Queen's University Belfast 2 , BT9 7BL Belfast , UK
3. Global Health Institute, School of Life Sciences, EPFL 3 , CH-1015 Lausanne , Switzerland