1. University of Virginia 1 Department of Cell Biology , , Charlottesville, VA 22908 , USA
2. University of Virginia 2 Department of Biomedical Engineering , , Charlottesville, VA 22903 , USA
3. University of Florida 3 Department of Medicine , , Gainesville, FL 32610 , USA
4. The Biocomplexity Institute, Indiana University 4 Department of Intelligent Systems Engineering and , Bloomington, IN 47408 , USA