1. Department of Ecology and Evolutionary Biology, University of Toronto, 25 Willcocks St., Toronto, ON, M5S 3B2, Canada
2. Department of Integrative Biology, University of Guelph, Guelph, Ontario, N1G 2W1, Canada
3. School of the Environment, University of Toronto, 33 Willcocks Street, Toronto, ON, M5S 3E8, Canada