1. Iowa State University 1 Department of Ecology, Evolution, and Organismal Biology , , Ames, IA 50011 , USA
2. University of North Carolina at Chapel Hill 2 Department of Biology , , Chapel Hill, NC 27599 , USA
3. Yale University 3 Department of Ecology and Evolutionary Biology , , New Haven, CT 06511 , USA
4. Faculty of Life Sciences, School of Zoology, Tel Aviv University 4 , Tel Aviv 6997801 , Israel