1. Department of Anthropology, University of Arkansas, Fayetteville, AR 72701, USA
2. Center for Functional Anatomy and Evolution, The Johns Hopkins University School of Medicine, Baltimore, MD, 21205, USA
3. Department of Medical Anatomical Sciences, Western University of Health Sciences, Pomona, CA, 91766, USA
4. Departments of Biological Sciences, Aerospace & Mechanical Engineering, and Anthropology, University of Notre Dame, Notre Dame, IN 46556, USA