1. Graduate School of Pharmaceutical Sciences, IRCN, The University of Tokyo, Tokyo 113-0033, Japan
2. Department of Cell Biology and Neuroscience, Juntendo University School of Medicine, Tokyo 113-8421, Japan
3. Laboratory for Cell Dynamics Observation, Center for Biosystems Dynamics Research (BDR), RIKEN, Osaka 565-0874, Japan
4. Department of Physics, Universal Biology Institute (UBI), and the International Research Center for Neurointelligence (WPI-IRCN), The University of Tokyo, Tokyo 113-0033, Japan
5. Division of Organelle Homeostasis, Medical Institute of Bioregulation, Kyushu University, Fukuoka 812-8582, Japan