1. Department of Pathology, University of Cambridge, Tennis Court Road, Cambridge CB2 1QP, UK
2. Department of Veterinary Medicine, University of Cambridge, Madingley Road, Cambridge CB3 0ES, UK
3. Department of Pharmacology, University of Cambridge, Tennis Court Road, Cambridge CB2 1PD, UK
4. School of Cellular and Molecular Medicine, University of Bristol, University Walk, Bristol BS8 1TD, UK