1. Department of Experimental Genome Research, Research Institute for Microbial Diseases, Osaka University 1 , Osaka 5650871 , Japan
2. Regulation of Host Defense Team, Center for Infectious Disease Education and Research, Osaka University 2 , Osaka 5650871 , Japan
3. Laboratory of Reproductive Systems Biology, The Institute of Medical Science, The University of Tokyo 3 , Tokyo 108-8639 , Japan