1. SOKENDAI (The Graduate University for Advanced Studies) 1 Department of Evolutionary Studies of Biosystems , , Shonan Village, Hayama, Kanagawa 240-0193 , Japan
2. NODAI Genome Research Center, Tokyo University of Agriculture 2 , 1-1-1 Sakuragaoka, Setagaya, Tokyo 156-8502 , Japan
3. Tokyo University of Agriculture 3 Department of Bioscience , , 1-1-1 Sakuragaoka, Setagaya, Tokyo 156-8502 , Japan