1. Department of Ecology, Evolution and Marine Biology, University of California, Santa Barbara, CA 93106, USA
2. Fish Ecology and Conservation Physiology Laboratory, Department of Biology and Institute of Environmental and Interdisciplinary Science, Carleton University, Ottawa, ON K1S 5B6, Canada
3. Fisheries and Oceans Canada, Cooperative Resource Management Institute, School of Resource and Environmental Management, Simon Fraser University, Burnaby, BC, Canada, V5A 1S6
4. Department of Psychology, Neuroscience & Behaviour, McMaster University, Hamilton Ontario, Canada L8S 4K1
5. Department of Zoology, University of British Columbia, Vancouver, BC, Canada, V6T 1Z4
6. Department of Zoology and Faculty of Land and Food Systems, University of British Columbia, Vancouver, BC, Canada, V6T 1Z4
7. Pacific Salmon Ecology and Conservation Laboratory, Department of Forest and Conservation Sciences, University of British Columbia, Vancouver, BC, Canada, V6T 1Z4